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Anything the expedites the arduous process of figuring out "Has anyone said anything directed at me since I last checked?" on a forum is huge.
Reddit has orange envelopes. Imagine how painful it'd be without them.
@Username mentioning is definitely a killer feature that comes to mind for a forum.
vBulletin's state of the art is for users to subscribe to topics they care about so they can scan each one to see if anyone quoted or replied to them. Newer forums like Discourse (meta.discourse.org) specifically improve in this area.
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